Large language models instantiate evolutionarily robust strategies of cooperation

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Authors: Saptarshi Pal, Abhishek Mallela, Lenz Pracher, Chiyu Wei, Feng Fu, Santiago D. Schnell, Martin A Nowak
Cited by: 1
References: 56

Abstract

Large language models (LLMs) are increasingly deployed to support human decision-making. This use of LLMs has concerning implications, especially when their prescriptions affect the welfare of others. To gauge how LLMs make social decisions, we explore whether five leading models produce sensible strategies in the repeated prisoner's dilemma, which is the main metaphor of reciprocal cooperation. First, we measure the propensity of LLMs to cooperate in a neutral setting, without using language reminiscent of how this game is usually presented. We record to what extent LLMs implement Nash equilibria or other well-known strategy classes. Thereafter, we explore how LLMs adapt their strategies to changes in parameter values. We vary the game's stopping probability, the payoff values, and if the total number of rounds is commonly known. We also study the effect of different framings. In each case, we test whether the adaptations of the LLMs are in line with basic intuition, theoretical predictions of evolutionary game theory, and experimental evidence of human participants. While all LLMs perform well in many of the tasks, none of them exhibit full consistency over all tasks. We also conduct tournaments between the inferred LLM strategies and study direct interaction between LLMs in games over ten rounds with known or unknown last round. Our experiments shed light on how current LLMs instantiate reciprocal cooperation.

Fields of Study & Subjects

Language and cultural evolution Evolutionary Game Theory and Cooperation Psychology of Moral and Emotional Judgment

Publication details

Journal / source
PNAS Nexus
Publisher
Oxford University Press (OUP)
Volume
5
Issue
6
Pages
pgag210–pgag210
Publication date
30 May 2026
Language
EN