On the free-living bryozo-an fauna from the Miocene Gram Formation (Denmark), with description of two new species, a new genus and a new suborder of Cheilostomata.

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Authors: Eckart Håkansson, Sten L. Jakobsen, Antonietta Rosso, Erik Thomsen
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Abstract

The free-living part of the restricted bryozoan fauna in the Late Miocene (Tortonian) Gram Formation, Denmark is described. Four species are recorded: Cupuladria aff. biporosa, Lunulites voigti, Reussirella germinatio n. sp. and Gramipora laxevincta n. gen., n. sp. The new suborder Cupuladriina nov. is introduced. The concepts of ‘free-living’ vs. ‘free-lying’ colonies are discussed briefly, with three categories of ‘free-living’ colonies considered, all exemplified by the four free-living taxa in the Gram fauna.

Fields of Study & Subjects

Paleontology and Stratigraphy of Fossils Paleontology and Evolutionary Biology Evolution and Paleontology Studies

Publication details

Journal / source
Bulletin of the Geological Society of Denmark
Volume
75
Pages
11–32
Publication date
12 January 2026
Language
EN