Contribution to the knowledge of the tribe Helopini Latreille, 1802 (Coleoptera: Tenebrionidae) of Iran

Two new species of darkling beetles of the genera Euboeus Boieldieu, 1865 and Eustenomacidius Nabozhenko, 2006 are described and illustrated: Eustenomacidius (Caucasohelops) rajabii Eshraghi Mofrad et Nabozhenko, sp. nov. (Alborz Mts: Ardabil Province) and Euboeus (Pelorinus) danae Eshraghi Mofrad et Nabozhenko, sp. nov. (Alborz Mts: Qazvin Province). The rank of one taxon is elevated from subspecies to species: Eustenomacidius (Caucasohelops) araxi Nabozhenko, 2006, stat. nov. (from E. svetlanae araxi). The following subgeneric synonymy is proposed: Caucasohelops Nabozhenko, 2006 = Zophondrus Nabozhenko, 2014, syn. nov. One new combination is established: Eustenomacidius (Caucasohelops) iranensis (Nabozhenko, 2014), comb. nov. (from the genus Zophohelops Reitter, 1902, the subgenus Zophondrus). Eustenomacidius (Caucasohelops) svetlanae Nabozhenko, 2006, and Adelphinus ordubadensis Reitter, 1890 are recorded from Iran for the first time. Following the present revision, the number of taxa of Helopini in Iran is increased to 61 species and three subspecies. The distribution of several previously described species in the tribe Helopini is discussed. Information on the range of Nalassus lineatus (Allard, 1876) is corrected. This species occurs only in the Hyrcanian forest in south-east Azerbaijan and Iran, while its occurrence in the Western Caucasus is excluded. The genus Turkmenohelops G.S. Medvedev, 1987 was erroneously reported from Iran and thus is excluded from its fauna.