Boulenophrys youran Xiao, Shi & Mo 2026, sp. nov.
Abstract
Boulenophrys youran Xiao, Shi & Mo sp. nov. Figs 5, 6, 7, 8 Type materials. Holotype. • HUNL XB 25061601, adult male, collected by Bei Xiao, Leqiang Zhu, Hongyu Wang on 16 June 2025 from Nanshan National Park (26°9'23.59"N, 110°10'47.32"E, ca 1 040 m a. s. l.), Chengbu Miao Autonomous County, Shaoyang City, Hunan Province, China, (Fig. 5). Paratypes. (n = 11) • HUNL XB 25061602 and HUNL XB 25061603, two adult males, collected from the same place as holotype. • HUNL XB 24052601 - HUNL XB 24052603, three males, collected on 26 May 2024 by Bei Xiao, Leqiang Zhu, and Ayinuer Maimaiti; • HUNL LH 23051201, adult male, and HUNL LH 23051202, adult female, collected from Yaorenping Village (26°14'50.43"N, 110°30'34.74"E, 1 090 m a. s. l.) on 12 May 2023 by Hui Li and Leqiang Zhu; • HUNL DB 24042601 and HUNL DB 24042602, two adult males, HUNL DB 24042603, adult female, HUNL DB 23092403, juvenile male, collected from Fanshuijiang Village, Xinning County, Hunan province (26°23'4.17"N, 111°1'16.86"E, 1 206 m a. s. l.) on 26 April 2024 and 24 September 2023 by Dongbo Fei, Changle Liao. Etymology. The specific epithet youran is a noun in apposition derived from the Chinese word “ 悠然 ” (yōu rán), meaning tranquil or composed. It references the verse “ 采菊东篱下, 悠然见南山 ” from the classical poem Drinking Wine No. 5 by Tao Yuanming (365–427 CE), which depicts a state of serene harmony with nature. The name thus alludes to the species type locality, Nanshan (南山), and evokes the pristine, undisturbed montane streams it inhabits. The English vernacular name “ Youran Horned Toad ” is here proposed, with the corresponding Chinese name “ 悠然角蟾 ” (Yōu Rán Jiǎo Chán). Diagnosis. This species can be distinguished from its congeners by the following combination of morphological characters: (1) Body size relatively large, SVL 45.3–51.1 mm in nine adult males, 55.6–58.2 mm in two adult females; (2) head wider than long, males HDL / HDW 0.69–0.83; (3) margin of tongue feebly notched posteriorly; (4) subarticular tubercles present on fingers I, relative finger lengths II IV> II> I; fingertips slightly dilated and rounded, without lateral fringes, subarticular tubercles only on fingers I, absent on fingers II – IV, replaced by low callous dermal ridges; inner metacarpal tubercle prominently enlarged, outer metacarpal tubercle smaller and flattened. Dense moderate black conical nuptial spines present on inner two fingers. Hindlimbs slender, length approximately 1.7 times of SVL, tibia length 0.56 of SVL, thigh length shorter than tibia length, but greater than foot length; heels overlapping when flexed hindlimbs are held at right angles to the body axis; tibiotarsal articulation exceeding snout tip when leg stretched forward; relative toes length: I
Fields of Study & Subjects
Diatoms and Algae Research
Genomics and Phylogenetic Studies
Bryophyte Studies and Records
Publication details
- Journal / source
- Zenodo (CERN European Organization for Nuclear Research)
- Publication date
- 5 January 2026