Netelia (Paropheltes) savchenkoi Tolkanitz 1981

Netelia (Paropheltes) savchenkoi Tolkanitz, 1981 Figs 6 C, 10, 11, 17 H – K Netelia (Netelia) savchenkoi Tolkanitz, 1981 b: 35. Netelia (Paropheltes) savchenkoi – Konishi 1996 a: 183. Diagnosis. This species can be distinguished from other species of subgenus Paropheltes in having the combination of the following characters: occipital carina complete in most specimens, absent dorsally in some specimens; scutellum with lateral carina present on anterior 0.8 to 0.9; gonostyle with submarginal spine present; pad rounded rectangular and projected dorso-basally, connected with conjunctiva ventro-apically; brace broadened basally and narrow apically. Type material. Holotype. Russia • 1 ♂; “ Приморский край, Кедровая падь ” [Primorsky Krai, Kedrovaya Pad]; 21.VI.1976; “ Савченко ” [Savchenko] leg.; light trap; Voucher: SIZU (Fig. 11). Non-type material. Taiwan – Nantou County • 1 ♂; Xingyi Township, Tatajia, Tai- 18 th highway 100.5 k, 23.480555°N, 120.852519°E; 16.VI.2020; light trap; C. - L. Huang leg.; GenBank: PV 910511 (COI); Sample ID: CHP 01326; Voucher: NMNS ENT 9068-2. 1 ♂; Ren’ai Township, Meifeng; 24.IV.1991; UV light trap; C. S. Sun leg.; Voucher: NMNS ENT 818-21. 3 ♂♂; ibid, Tayuling, 820 forest road (Fogeste); UV light trap; 3.IX.2008; H. H. Lin leg.; Voucher: NMNS ENT 6070-580, 1199–1200. 2 ♂♂; ibid, Tsuifeng; alt. 2300 m; Malaise trap; X.1985; K. S. Lin leg.; Voucher: TARI Net 32, 41. – Hualien County • 1 ♂; Xiulin [Hsiulin] Township; Kuan-yun Hostel; alt. 2335 m; 27–28.V.2008; UV light trap; Liang, Chen, Fu leg.; Voucher: NMNS ENT 5921-1017. – Chiayi County • 1 ♂; Alishan Township, Alishan, 23.47666°N, 120.71583°E; 28.IV.2023; light trap; C. - Y. Juang leg.; GenBank: PV 910510 (COI); Sample ID: CHP 01278; Voucher: TARI. Redescription based on Taiwanese specimens. Measurements were based on 9 males. Male. Head (Figs 10 A – C, 11 C – E): vertex matt, weakly and finely coriaceous; frons, face and clypeus matt and coriaceous with setae; HW / HL = 1.6–2.2 (1.9 ± 0.18), lateral ocelli almost touching eyes, OD = 0.2–0.3 (0.26 ± 0.01) mm, POL / OD = 0.3–0.5 (0.38 ± 0.06); occipital carina complete, sometimes absent dorsally; face FW / FH = 1.0–1.2 (1.09 ± 0.06); clypeus CLW / CLH = 2.0–2.4 (2.17 ± 0.13), ventral margin weakly concave; malar space with MSL / BMW = 0.2–0.6 (0.33 ± 0.10); gena GOI = 2.8–4.0 (3.36 ± 0.43); flagellum with 45–48 segments, 1 st flagellomere 1.2–1.4 (1.27 ± 0.09) × as long as 2 nd flagellomere, 3.9–5.8 (4.58 ± 0.57) × as long as wide. Mesosoma (Figs 10 D, E, 11 E, F): pronotum polished, minutely punctate dorsally and rugose medially; mesoscutum and scutellum matt, evenly and minutely punctate; mesopleuron polished and minutely punctate; mesoscutum MSSL / MSSW = 1.1–1.4 (1.27 ± 0.08), with notaulus present on about anterior 0.4; scutellum SCL / SCW = 1.1–1.7 (1.42 ± 0.19), with lateral carina present on anterior 0.8 to 0.9; propodeum evenly and minutely punctate on anterior half, with dense and transverse striae posteriorly, sublateral crest normal, not connected dorsally. Wings (Figs 10 F, 11 B): fore wing length 8.5–10.2 (9.42 ± 0.60) mm; 3 rs-m present, areolet medium-sized and anteriorly stalked; 2 rs-m and abscissa between 2 rs-m and 2 m-cu in acute angle; RMI = 0.8–1.5 (1.04 ± 0.20); ramulus absent; 1 cu-a straight and vertical, distad to M & RS with BNI = 0.1–0.4 (0.24 ± 0.08); cell AA with setal band narrow and sparse on about proximal 0.3–0.5; AA almost straight; hind wing with distal 6–7 hamuli; NI = 0.3–0.4 (0.33 ± 0.05). Legs: fore tibial spur 0.3–0.4 (0.31 ± 0.06) × as long as 1 st tarsomere; fore 1 st tarsomere 14.0–19.3 (17.36 ± 1.72) × as long as wide, 1.7–1.9 (1.77 ± 0.08) × as long as 2 nd; fore tarsal claw with 16–19 pectens; hind 1 st tarsomere 11.3–16.1 (14.24 ± 1.53) × as long as wide, 1.8–2.1 (1.97 ± 0.11) × as long as 2 nd; hind tarsal claw normal, with 12–15 pectens. Metasoma (Figs 17 H – K, 11 H – J): T 1 4.0–5.2 (4.41 ± 0.41) × as long as posterior width, 1.4–1.7 (1.56 ± 0.09) × as long as T 2; abdominal sternum IX HPW / HPL = 0.8–1.1 (0.99 ± 0.09), posteriorly rounded; gonostyle rounded and weakly tapered apically, with submarginal spine present on about dorso-apical 0.2, GSWA / GSL = 0.3 (0.31 ± 0.02), GSWB / GSL = 0.5–0.6 (0.56 ± 0.05), GSWB / GSWA = 1.7–1.9 (1.83 ± 0.05), pad rounded rectangular and projected dorso-basally, connected with conjunctiva ventro-apically; brace broad basally and narrow apically; gonossiculus short, broad apically and pointed ventrally; basivolsella with patch of sparse and short seta; penisvalva PVL / GSL = 0.5–0.7 (0.58 ± 0.04), volviceps with dorsal margin about 0.9 length of ventral margin, valvura strongly and dorsally curved. Coloration (Figs 6 C, 10, 11): head yellowish brown, except face yellowish white and dorsal portion of occiput sometimes brown; mesosoma yellowish brown, with mesoscutum and dorsal surface of propodeum brown, speculum and anterior corner of mesopleuron yellowish white; wings hyaline, with wing vein yellowish brown and pterostigma pale yellowish brown; legs yellowish brown, except hind coxa brown, hind tarsus yellowish white to yellow; metasoma yellowish brown. Female. Taiwanese specimens not examined. Described in Tolkanitz (1981 b) and Konishi (1996 a). Distribution. Russia (Promorskii Krai [type locality] (Tolkanitz 1981 b)), Japan (Honshu, Shikoku, Kyushu (Konishi 1996 a)), Taiwan (new record: Nantou, Hualien, Chiayi). Bionomics. This species was collected from the mountains between the elevation 1320–2400 m in Taiwan. This new record extends the distribution of an eastern Palearctic species to the mountain areas of a subtropical region. Remarks. Despite minor variations in general morphology and male genitalia observed between the Taiwanese and eastern Palearctic specimens, these differences are considered intraspecific variation. For instance, the lateral carina extends beyond the anterior 0.8 in Russian and Taiwanese specimens but only reaches 0.8 in Japanese specimens (Konishi 1996 a), and the brace is narrower in Taiwanese specimens but broader in Japanese and Russian specimens (Tolkanitz 1981 b; Konishi 1996 a). This species can be assigned to the N. (Paro.) barberi species group based on the brownish mesosoma without any distinctive yellow marking and lateral carina of scutellum extending beyond 0.7 of its length.