Deleproctophylla australis

Deleproctophylla australis (Fabricius, 1787) Figs 2 A, 2 B, 4 A – D, 5 A, 5 B, 6–8 Ascalaphus australis Fabricius, 1787: 250 (type locality: “ Europa australiore ” = southern Europe; holotype in NHMD). Gmelin 1788: 2645 (Myrmeleon). Villers 1789: 62 (Myrmeleon). Olivier 1790: 245 (Ascalaphus). Fabricius 1793: 96 (Ascalaphus). Turton 1800: 405 (Myrmeleon). Angelini 1827: 468 (Ascalaphus). Lefèbvre 1842: 6 (Deleproctophylla). Rambur 1842: 351 (Theleproctophylla). Schneider 1845: 341 (Theleproctophylla). Walker 1853: 416 (Ascalaphus). Hagen 1860: 53 (Theleproctophylla). Stein 1863: 420 (Ascalaphus). Hagen 1866: 402 (Deleproctophylla). Costa 1866: 14 (Theleproctophylla). McLachlan 1873: 260 (Theleproctophylla). Navás 1905: 547 (Theleproctophylla). Navás 1911: 92 (Theleproctophylla). Navás 1912 a: 124 (Theleproctophylla). Navás 1912 b: 34 (Theleproctophylla). Lacroix 1913: 100 (Theleproctophylla). Kimmins 1930: 188 (Theleproctophylla). Mosely 1932: 182 (Theleproctophylla). Táborský 1936: 164 (Deleproctophylla). Capra 1936: 213 (Theleproctophylla). Capra 1937: 58 (Theleproctophylla). Zimsen 1964: 612 (Ascalaphus). Zelený 1964: 326 (Theleproctophylla). Popov 1977: 276 (Deleproctophylla). H. Aspöck et al. 1980: 315 (Deleproctophylla). Devetak 1992: 111 (Deleproctophylla). Zakharenko and Krivokhatsky 1993: 73 (Deleproctophylla). Bernardi et al. 1995: 11 (Deleproctophylla). Devetak 1995: 194 (Deleproctophylla). Letardi and Pantaleoni 1996: 299 (Deleproctophylla). H. Aspöck and Hölzel 1996: 78 (Deleproctophylla). Pantaleoni and Letardi 1998: 40 (Deleproctophylla). H. Aspöck et al. 2001: 301 (Deleproctophylla). Whittington 2002: 377 (Deleproctophylla). Popov 2004: 231 (Deleproctophylla). Canbulat 2007: 36 (Deleproctophylla). Dobosz and Ábrahám 2007: 24 (Deleproctophylla). Devetak and Janžekovič 2012: 183 (Deleproctophylla). Colombo et al. 2012: 125 (Deleproctophylla). Badano and Pantaleoni 2014: 301 (larva, Deleproctophylla). Letardi 2016: 194 (Deleproctophylla). Aistleitner 2019: 123 (Deleproctophylla). Jones 2019: 3 (Deleproctophylla). Poggi 2024: 9 (Deleproctophylla). Theleproctophylla australis var. impar Navás, 1923: 154 (type locality: Greece: Thessaloniki; holotype in MNHN but lost at present). Examined material. • Croatia: Dalmatien, Hvar, 17. VII. 1962, R. Löberbauer, 1 ♀ (HUAC); • Croatia: Dalmatien-Split, Küste, 22. VI. 1953, H. Hamann, 1 ♂ 1 ♀ (HUAC); • Croatia: Ribnica, Otavice 10 km N v. Titov Velea, 28. VII. 1965, 1 ♂ 1 ♀ (RMNH); Croatia: • Rovinj, 15. V. 1953, 1 ♂ (HUAC); • Croatia: Ninski Stanovi, 40 m, 14. 07. 2004, L. Ábrahám, 31 ♂♂ 34 ♀♀ (SCM); • Croatia: Zadar, Benkovac, 11. 07. 2002, D. Szalóki, 3 ♀♀ (SCM); • Croatia: Dalmacija, Gor. Poličnik, 12 km E of Zadar, 20. 06. 2007, leg. Zs. Józan, 1 ♀ (SCM); • Croatia: Dalmacija, Primošten-Bilo, 09. 06. 2003, Nikola Rahmé 1 ♂ (SCM); • Croatia: Vinodol Obc., Crikvenica, Selce, 180 m, 01. 07. 2001, K. Székely 1 ♂ (SCM); • Croatia: Ninski Stanovi, 40 m, 14. 07. 2004, L. Ábrahám, 3 ♂♂ 3 ♀♀ (USMB); • France: Corse, Calvi, 13–27. VII. 1971, Fam. V. Oorschot, 1 ♀ (RMNH); • France: Corse, Belgodère, 350 m, 14. VII. 1971, Fam. V. Oorschot, 1 ♀ (RMNH); • France: Corse, Belvedere, VII. 1973, H. M. Steiner, 3 ♂ 1 ♀ (HUAC); • France: Corsica, Valle de Restino, 02. 06. 2000, leg. L. Nádai, 1 ♀ (SCM); • France: Haute-Corse, Castifao, 25. 06. 2002, leg. J. Bard, 1 ♀ (SCM); • Greece: Ahaia, Michas, 1300 m, 38°00'34"N, 21°53'00"E, 7. VII. 2008, Dils J. and Faes J., 1 ♀ (RMNH); • Greece: Chalcide, Tristinika, 20. VI. 2015, 1 ♂ (MDC); • Greece: Cos, Tigaki, 3–10. VII. 1978, M. C. and G. Kruseman, 2 ♀ (RMNH); • Greece: Georgioupolis, 35°21'N, 24°16'E, H. u. L. Hölzel, 24 ♂ 19 ♀ (NHMW); • Greece: Georgioupolis, 35°21'N, 24°16'E, H. u. L. Hölzel, 1 ♂ 1 ♀ (HUAC); • Greece: Elis, Andritsaina, 1000 m, 29. VII. 1978, L. Willemse and J. Tilmans, 1 ♂ (RMNH); • Greece: Ioannina, Mesovouni, 650 m, 39°56'18.8"N, 20°38'12.4"E, 28. VII. 2006, T., W. and J. Garrevoet, 2 ♀ (RMNH); • Greece: Lesbos, Sigri, 6–8. VI. 1988, J. P. Duffels, 1 ♂ 1 ♀ (RMNH); • Greece: Loutraki-Perachora, 38°N, 23°E, 30. V. 1969, Rausch, 1 ♀ (HUAC); • Greece: Pierias, Litochoro, 5. VII. 1976, F. and L. Willemse and J. Tilmans, 1 ♀ (RMNH); • Greece, Creta Nord, Hersonissos, 35°18'36.3"N, 25°23'53.9"E, 15. V. 2024, J. Vetrovec, 1 ♂ (DB); • Greece: Creta, Lasithi, Xerokampos, 2. VI. 1972, M. C. and G. Kruseman, 1 ♂ 2 ♀ (RMNH); • Greece: Kardaras, 800 m, 37°39'22.9"N, 22°18'34.2"E, 7. VII. 2008, Dils J. and Faes J., 1 ♂ (RMNH); • Greece: Tripolis-Davia 3–8 km NW of Tripolis, 800 m, 13. VII. 1975, J. P. Duffels, 1 ♀ (RMNH); • Greece: Leptokaria, 28. 06. 1992, Balásházy, 1 ♀ (SCM); • Greece: Sithonia, Sarti, 0–50 m, 02–09.07. 1997, D. Szalóki, 1 ♂ 4 ♀♀ (SCM); • Greece: Sithonia, Sarti, 0–50 m, 02–09.07. 1997, D. Szalóki, 1 ♀ (USMB); • Greece: Argolída, SE of Néa Epídavros, 37°40'N, 23°09'E, 120 m, 09. 06. 2007, R. Królik, 1 ♂ (USMB); • Greece: Corfu, Acharavi [CK 90], xerothermic meadow, 06. 07. 2011, W. Żyła, 3 ♂♂ (USMB); • Greece: Corfu, Acharavi [CK 90], xerothermic meadow, 07. 07. 2011, W. Żyła, 1 ♂ (USMB); • Greece: Chalkidiki, Paliouri env., 39°56'31"N, 23°39'29"E, SE of Kassandria, 06–11.06. 2013, M. Šnižek, 2 ♂♂ (USMB); • Greece: Chalkidiki, Paliouri env., 39°56'31"N, 23°39'29"E, SE of Kassandria, 12–14.06. 2013, M. Šnižek, 3 ♂♂ (USMB); • Greece: Thessaly, Kokkino Nero, 39°49'N, 22°47'E, environs, 17–29.05. 2015, A. Skiba, 1 ♂ (USMB); • Greece: Chalkidiki, E of Paliouri, 39°56'N, 23°41'E, Kassandria, 11. 06. 2022, M. Šnižek, 1 ♂ (USMB); • Greece: Peloponnese, Loutra Killinis, 37°51'12"N, 21°6'37"E, dunes and grass, 19. 06. 2023, W. Żyła, 1 ♀ (USMB); • Greece: Peloponnese, Messenia, SE of Kalamia, 37°52'13.5"N, 21°07'08.7"E, olive garden, 13. 06. 2024, W. Żyła, 2 ♀♀ (USMB); • Greece: Peloponnese, Messenia, SE of Kalamia, 37°52'13.5"N, 21°07'08.7"E, olive garden, 14. 06. 2024, W. Żyła, 1 ♂ 1 ♀ (in absolute alcohol) (USMB); • Italy: Basilicata, 10 km uoru, Potenza, 700 m, 40°35'N, 15°45'E, 1 ♂ (HUAC); • Italy: Calabria, Crotone, Catanzaro, 30–100 m, 38°55'N, 16°55'E, 4. VIII, 1 ♂ (HUAC); • Italy: Prov. Messina, Lipari, Isole, Stromboli, San Vincenzo, 17. VII. 1995, Alain Olivier, 1 ♀ (RMNH); • Italy: Tuscany, Grosseto, Nomadelfia (GR), 42°50'23"N, 11°08'50"E, 29. VI. 2024, Davide Badano and Yuchen Zheng, 2 ♂ 2 ♀ (ZCAU); • same locality • 1 ♂ 3 ♀ (DB); • Italy: Tuscany, Grosseto, Roselle (GR), 42°49'34"N, 11°10'05"E, 29. VI. 2024, Davide Badano and Yuchen Zheng, 1 ♂ 2 ♀ (ZCAU); • same locality data • 1 ♀ (DB); • Italy: Tuscany, Grosseto, Parco Nazionale della Maremma, Marina di Alberese (GR), 42°39'51"N, 11°02'25"E 20. VI. 2025, Davide Badano leg., 1 ♂ 2 ♀ (DB); • Italy: Lazio, Roma (RM), Via Appia Antica, 11. VII. 1937, L. Barbera, 1 ♀ (MSNG); • Italy: Nomadelfia (GR), 42°50'23.2"N, 11°08'51.4"E, 30. VI. 2024, D. Badano and Y. Zheng, (DB); • Italy: Puglia, Bari, Masseria Pellicciari, 450 m, 9. VII. 2000, Fontana P., Kleukers R. and Cogo A., 1 ♀ (RMNH); • Italy: Puglia, Castel del Monte, 450 m, 41°05'20.9"N, 16°16'16.3"E, 2. VII. 2001, T. and W. Garrevoet, 1 ♀ (RMNH); • Italy: Prov. Salerno, Pioppi, Cost Cilentana, 14. VII. 1971, Holzinger, 1 ♀ (HUAC); • Italy: Sardegna, Marrubiu (OR), Sant’Anna, stazione e dintorni, 1. VI. 2003, C. Meloni, 1 same locality, ♂ (MSNG); • same locality, 2. VI. 2006, 1 ♀ (MSNG); • same locality, 26. VI. 2009, C. Meloni, 3 ♂ 12 ♀ (MSNG); • Italy: Sardegna, Alghero (SS), Lu Cantar, 40°32'25.3"N, 8°19'29.7"E, 12. VII. 2012, D. Badano, 3 ♂ (DB); • Italy: Sardegna, Alghero (SS), Monte Doglia, 40°36'20.1"N, 8°15'06.0"E, 7. VII. 2012, D. Badano, 3 ♀ (DB); • Italy: Sardegna, Monti (OT), 2. VII. 2011, D. Badano, 2 ♀ (DB); • Italy: Sicilia, Sud di Castrofilippo (AG), m 450, 4. VI. 2015, E. Gallo, 1 ♀ (MSNG); • Italy: Sicilia, Caltanissetta (CL), strada Enna, bivio Gela, 8. VII. 1954, Adalberto Parvis, 2 ♂ 3 ♀ (MSNG); • Italy: Sicilia, Peloritani, 10 km SW Barcellona, 100 m, 38.06°N, 15.07°E, 25–26.1974, 2 ♂ 2 ♀ (HUAC); • Italy: Sardinia, Siniscola, Su Tilio, 27. 06. 2004, V. Feik, 1 ♀ (NMPC); • Italy: Sardinia, “ Sardinien, Mti Gennargentu Mte, Spada ”, 1250 m, 17–21.06. 1968, Ströhle, 2 ♂♂ (SCM); • North Macedonia: Demir Kapija, Barovo, 27. VII. 1965, 1 ♀ (RMNH); • North Macedonia: Štip, 23. VII. 1965, 1 ♂ (RMNH); • Turkey: Aydin, Kusadasi, 11. VII. 1980, H. v. Oorschot, 1 ♀ (RMNH); • Turkey: Balıkesir, nahe Bezirci (zw. Edincik u. Bugdayli), 150 m, 40.15°N, 27.47°E, 22. VII. 1978, H. and U. Aspöck, H and R Rausch, P. Ressl, 3 ♂ 4 ♀ (HUAC); • Turkey: Edirne, 08–13.06. [19] 47, 1 ♂, Exp. N. Mus. ČRS (NMPC). Diagnostic characters. Metafemur with brown marking; forewing with brown marking below pterostigma (often absent in eastern populations); hindwing elongate, posterior margin curved; not angled; hindwing with large marking below pterostigma; male ectoproct with branch in apical half of branch, after mid-length; setae on branch only at tip; setae on branch and tip of ectoproct stout and curved. Redescription. Size (mm, based on 5 specimens). Head + thorax length 7.7–8.6; antenna length 15.1–2; fore wing length 21.7–22.9; fore wing width 6.4–7.4; hindwing length 17.1–18.7; hindwing width 5.0–5.7. Head. Vertex yellowish brown; setae dense, long, pale yellow (Fig. 6 C). Para-ocular band pale yellow. Frons yellowish brown; setae dense, long, pale yellow. Clypeus and labrum yellowish brown; setae long, pale (Fig. 6 B). Mandible yellowish brown, transitioning to dark brown apically. Labium and palp yellowish brown, concolorous with mandible. Antenna with pedicel with dense pale and dark setae, flagellomeres brown, nodes slightly darker; club brown, anterior face paler (Fig. 6 A). Thorax. Cervical sclerite yellowish brown. Pronotum brown; setae long, pale yellow. Mesonotum brown with yellowish brown markings: an anterior rounded pair, a median triangular pair, and a posterior rounded pair; mesoscutellum anteriorly dark brown, posteriorly yellowish brown. Mesonotum with long pale yellow setae. Metanotum la