Deleproctophylla gelini

Deleproctophylla gelini (Navás, 1919) Figs 3 A, 3 B, 15, 16 Theleproctophylla gelini Navás, 1919: 21 (type locality: Morocco: Safi; holotype in MNHN but lost at present). Navás 1930: 121 (Theleproctophylla). H. Aspöck and Hölzel 1996: 78 (Deleproctophylla). H. Aspöck et al. 2001: 302 (Deleproctophylla). Whittington 2002: 377 (Deleproctophylla). Ábrahám and Mészáros 2004: 333 (Deleproctophylla). Faucheux 2006: 204 (Deleproctophylla). Faucheux et al. 2012: 171 (Deleproctophylla). Ábrahám 2017: 126 (Deleproctophylla). Examined material. • Morocco: Agadir, 27. V – 6. VI. 1973, C. and A. Jeekel, 2 ♀ (RMNH); • Morocco: Bou Tazzert, Pres Mogadoa, 1928, R. Benoist. 2 ♂, 1 ♀, 2 ex. (MNHN); • Morocco: Tamri, 30°42'31.9"N, 09°51'26.1"W, 15 m, 30. 06. 2008, L. Ábrahám, L. Bognár and L. Nagy, 5 ♂♂ 1 ♀ (SCM); • Morocco: Tamri, 30°42'31.9"N, 09°51'26.1"W, 15 m, 30. 06. 2008, L. Ábrahám, L. Bognár and L. Nagy, 1 ♂ 1 ♀ (USMB); • Morocco: Atlantic Plain, 4 km W Agoudi n’Aït Amar 30°42'31.3"N, 9°51'26.2"W, 10 m, 27. 06. 2024, W. Szczepański, 4 ♂♂ 2 ♀♀ (USMB); Diagnostic characters. Vertex as long as wide; metafemur without brown marking; forewing extensively marked; hindwing broad, posterior margin curved; forewing with complex pattern: brown marking at base, subcostal area brown, brown marking at RP bifurcation, and large brown marking from pterostigma to first branch of RP; hindwing with a similar pattern: brown marking at base, subcostal area brown, brown marking at RP bifurcation, and large brown marking from pterostigma to posterior margin; male ectoproct bent inward, with branch at mid-length and protruding dorsally; setae on branch only at tip; setae on tip of ectoproct robust and relatively long; male with gx 11 straight, broad, incapsulating gx 9; gx 9 without apical fold; female ectoproct long; gx 8 and gp 8 not prominent. Redescription. Size (mm, based on 3 specimens). Head + thorax 7.4–9; antenna length 14.9–16.9; fore wing length 21–24.9; fore wing 6.2–7.3; hindwing 16.1–20.5; hindwing width 5.2–6.1. Head. Vertex nearly as long as wide, yellowish brown; setae dense, long, pale yellow (Fig. 15 C). Para-ocular band pale yellow. Frons yellowish brown; setae dense, long, pale yellow. Clypeus and labrum yellowish brown; setae long, pale (Fig. 15 B). Mandible yellowish brown, dark brown apically. Labium and palp yellowish brown. Antenna with pedicel with dense pale and dark setae, flagellomeres pale brown, nodes dark brown; club dark brown, paler anteriorly (Fig. 15 A). Thorax. Cervical sclerite yellowish brown. Pronotum brown; setae long, pale yellow. Mesonotum yellowish brown gradually darker medially and along sclerites margins. Metanotum yellowish brown, medially darker; setae pale yellow (Fig. 15 C). Pleurae yellowish brown; setae pale yellow (Fig. 15 D). Legs. Pro- and mesothoracic legs yellowish brown, including tarsi. Metathoracic leg yellowish brown, unmarked. Setae of coxae pale yellow setae; setae of femora and tibiae thin and pale yellow dorsally, thicker and blackish ventrally; tibial spurs narrow; setae of tarsi blackish (Fig. 15 D). Wings. Forewing broad, apex rounded. Pterostigma with ~ 5 forked and unforked brown veinlets; membrane brown. Presectoral area with five crossveins. RP with ~ 5 branches. MA gently toward hind margin. MP - CuA area with ~ 3 irregular rows of cells. Anal area hind margin straight; axillary lobe slightly pronounced. Forewing venation largely pale brown. Forewing membrane largely hyaline with a complex brown pattern: base of wing brown, apical costal crossveins with brown shade at base, subcostal area brown, brown marking at RP branching, apical portion of wing with a large brown marking extending from subcostal area and pterostigma to first branch of RP (Figs 3 A, 3 B, 15 A). Hindwing broad, widest at RP fork; hind margin rounded, smoothly curved toward rounded apex. Presectoral area with three crossveins. RP with five branches. MA largely straight, slightly curved distally. MP - CuA area with ~ 2 irregular rows of cells. Hindwing venation pale brown, Hindwing membrane hyaline, with a complex brown pattern: base of wing brown, apical costal crossveins with brown shade at base, subcostal area brown, brown marking at RP branching, apical portion of wing with a large brown marking extending from subcostal area and pterostigma to posterior margin of wing (Figs 3 A, 3 B, 15 A). Abdomen. Tergites pale brown. Sternites pale brown, paler than tergites. Pleural membrane brown. Male genitalia. Ectoproct robust, strongly bent inward, with branch slightly beyond mid-length. Branch projecting dorsally, broad, and relatively thick, progressively widening at apex. Apex of ectoproct club-shaped expanding internally. Setae of ectoproct short, dark brown; setae on branch sparse, those on apical section, black, stout, and curved; setae on tip of ectoproct progressively denser caudally, black, stout, curved, and spine-like on the internal side (Fig. 16 A – C). Tergite 9 trapezoid in lateral view, with broad ventral projection; setae long and black, longer and denser on projection (Fig. 16 B, C). Sternite 9 without paired lateral lobes, being composed only by the median lobe; setae thin, spine-like lobe. Gx 11 arch-shaped, projecting caudally, broad and incapsulating gx 9, straight in lateral view; gp 11 lightly sclerotized, elongated, with a shallow median furrow, with minute setae; gx 9 widely set apart, strongly sclerotized, flattened, without dorsal fold (Fig. 16 F – J). Female genitalia. Ectoproct strongly elongated, projecting ventrally; setae long, black (Fig. 16 D, E). Tergite 9 sub-rectangular. Gx 9 wider than long; setae long. Tergite 8 subrectangular. Gx 8 not prominent, with long setae. Gp 8 not prominent; setae thin. Sternite 7 with sparse stout setae (Fig. 16 D, E). Distribution. Africa: Morocco (H. Aspöck et al. 2001; Ábrahám and Mészáros 2004; Oswald 2025). Comparative notes. Among all Deleproctophylla species, D. gelini is the easiest to identify due to its unique fore- and hindwing pattern (Ábrahám and Mészáros 2004) (Fig. 3 A, B). Additionally, this species differs from all congeners in both male and female genitalia, which are characterized by several distinctive characters (Ábrahám and Mészáros 2004) (Fig. 16). The male ectoproct is stout and distinctly bent inward and the branch arises dorso-caudally at mid-length. The branch stacks on the ectoproct, instead of being directed inward, as seen in all other species. Furthermore, the sternite 9 of the male lacks the lateral lobes found in other species (Fig. 16 A – C). The gx 9 + 11 complex is also different being, shaped like a capsule. The Gx 11 is broad, not curved downward, and encases the gx 9, which lacks the apical dorsal fold present in all other species (Fig. 16 F – J). The female of D. gelini is characterized by an elongated ectoproct projecting ventrally and reduced gx 8 (Fig. 16 D, E).