Fynbosia horingaria Englund, Staude & Sihvonen 2026, sp. nov.
Abstract
Fynbosia horingaria Englund, Staude & Sihvonen sp. nov. Type material. Holotype • ♂ (Figs 3 A, 3 C, 5 A, 6 A), South Africa: Calitzdorp, Swartberg, farm Zandrivier, 33.373°S, 21.330°E, 1200 m, 28 Apr. 2022, Mikael Englund leg., DNA sample ME 09, gen. prep. PS 2872, micro-CT scan MS 2, coll. Ditsong Museum of Natural History, Pretoria, South Africa. Paratypes (5 ♂♂) • ♂ (Fig. 3 E), South Africa: Western Cape, 40 km SE of Calitzdorp, farm Zandrivier, dry renosterveld – nama karoo 33.351°S, 21.317°E, 1200 m, 28–29 Apr. 2022, Mikael Englund, Pasi Sihvonen, Hermann Staude leg., DNA sample PS 1641, photo PMS_31002, coll. Englund, Finland; • ♂ (Fig. 3 F), South Africa: Western Cape, 40 km SE of Calitzdorp, farm Zandrivier, dry renosterveld – nama karoo 33.351°S, 21.317°E, 1200 m, 28–29 Apr. 2022, Mikael Englund, Pasi Sihvonen, Hermann Staude leg., DNA sample PS 1659, photo PMS _ 31003, micro-CT scan MS 184, coll. Sihvonen, Finland; • ♂ South Africa: Western Cape, Calitzdorp, Seweweekspoort, Aristata, riverine fynbos, 33.4057°S, 21.4017°E, 12 Apr. 2023, A. Hofmann & J. Meineke, coll. Staude, South Africa; • ♂ South Africa: Western Cape, Calitzdorp, Seweweekspoort, Aristata, riverine fynbos, 33.4057°S, 21.4017°E, 12 Apr. 2023, A. Hofmann & J. Meineke, coll. State Museum of Natural History Stuttgart, Germany. Diagnosis. Adult male F. horingaria sp. nov. distinguishable from F. unicaria sp. nov. based on the wing patterns and genitalia. Both fore- and hindwing medial lines evenly rounded, not sharply angled near costa as in F. unicaria sp. nov., best visible on the wing undersides. Discal spots prominent, blackish, triangular in forewings (Figs 3, 6; Key 1). Male genitalia weakly sclerotized, but distinguishable from F. unicaria sp. nov. by the broader valva and elongated saccus (Figs 6, 8; Key 2). Average mtCOI genetic distance to F. unicaria sp. nov. is 5 %. Description. Adults. Head: Frons and vertex covered with light brown scales, mixed with some scattered dark brown scales, a string of hair-like scales protruding at the posterior margin of head, compound eye naked. Labial palpi short, with just two segments (Fig. 5 B), length less than 1 / 5 of eye diameter, oriented ventrally, covered with yellowish ochreous scales; vom Rath’s organ cavity on the tip of the terminal palpomere. Proboscis vestigial, hardly visible between labial palpi, apparently defunct. Male antenna (Figs 3, 6, 7) bipectinate, flagella dorsally covered with ochraceous cells, flagellomeres light brown, except for the blackish swollen end of each flagellomere, ramus black, conspicuously long, length up to twice the diameter of compound eye, abundant long cilia covering the ramus. Female antenna filiform. Thorax: Covered with mixed light to dark brown scales. Spur pattern on legs 0–0 – 2. Abdomen: Segments covered with brown scales of varying tone, mottled with blackish scales. Some long hair-like scales extending distally from the proximal segments of the male abdomen. Bowl-shaped paired tympanal organs laterally in the segments 1 + 2, tip of ansa slightly swollen. Wings (Figs 3 A, 3 C, 5 A, 6 A): Forewing triangular, elongated, termen slightly convex from apex until midwing (M 3), tornus round, without distinct transition from termen to dorsum. Wingspan 18–21 mm. Forewing ground colour ranging from light to medium brown with several dark wavy transverse lines most pronounced at costa. Proximal parts of medial and subcostal areas lighter brown than rest of forewing. Basal area with hazel brown tone. Discal spot distinct, triangular, blackish. Three dark brown blotches in terminal area, largest situated medially, smaller ones towards apex and tornus. Ante- and postmedial lines dark, evenly rounding inwards near costa in both fore- and hindwings. Hind wing apex rather sharply angled, discal spot dark. Terminal area lighter with darker suffuse plots forming discontinuous wavy line. Markings on hindwings more distinct on underside. Genitalia: Male (Figs 6 B, 8 A – E): Genitalia generally weakly sclerotised with few diagnostic ornaments, except for uncus and gnathos. Valva wide, costa evenly rounded. Uncus rather well sclerotised, prominent with evenly downward hooking tip, juxta sclerotised with saddle-form upward hooking tip. Saccus elongated, aedeagus lacking cornuti. Female genitalia unknown. Immature stages unknown. Genetic data. GenBank accession numbers PV 988405 – PV 988410. Etymology. Horing (> Afrikaans = horn); pertaining to the large and conspicuous, plume-like antennae of the male.
Fields of Study & Subjects
Coleoptera Taxonomy and Distribution
Lepidoptera: Biology and Taxonomy
Hymenoptera taxonomy and phylogeny
Publication details
- Publication date
- 30 January 2026