Campanistylus lakhraensis Merle & Pacaud 2026, n. gen., n. sp.

Campanistylus lakhraensis Merle & Pacaud, n. gen., n. sp. (Figs 5; 6) urn:lsid:zoobank.org:act: 501A660A-F529-48F8-8D9D-D49A3571A5F5 TYPE MATERIAL. — Holotype. Pakistan • Lower Indus Basin, Sindh Province, Jamshoro district; Lakhra Dome, East of Lakhra village section (stn 2), lower Lakhra Formation; early Eocene (Ypresian); CPAG. RAN.I.95 (cast MNHN. F.A93729); H: 136.8 mm (c. 32 whorls), D: 40.1 mm; Fig. 6A, B. Paratypes. Pakistan • 1 spm; same data as the holotype; CPAG. RAN.I.102 (cast MNHN. F.A93726), H: 31.9 mm (shell incomplete, 6 whorls), D: 17.9 mm; Fig. 5D • 1 spm; idem; CPAG. RAN.I.103 (cast MNHN. F.A93725), H: 57.1 mm (shell incomplete, 8 whorls), D: 27.1 mm; Fig. 5C • 1 spm; idem; CPAG. RAN.I.104 (cast MNHN. F.A93727), H: 29.2 mm (shell incomplete, 7 whorls), D: 14 mm; Fig. 5B • 1 spm; idem; CPAG.RAN.I.105 (cast MNHN. F.A93728), H: 58.6 mm (shell incomplete, 8 whorls), D: 29.7 mm; Fig. 5A • 1 spm; same data as for the holotype; CPAG. RAN.I.96 (cast MNHN. F.A98131), H: 159.4 mm (c. 27 whorls), D: 43.3 mm; Fig. 6C • 1 spm; idem; CPAG.RAN.I.97 (cast MNHN. F.A98129), H: 114.9 mm (shell incomplete, 6 whorls), D: 43.8 mm • 1 spm; idem; CPAG.RAN.I.98 (cast MNHN. F.A93712), H: 91.4 mm (shell incomplete, 3 whorls), D: 46.1 mm; Fig. 6H, I • 1 spm; idem; CPAG.RAN.I.99 (cast MNHN. F.A98132), H: 115.4 mm (shell incomplete, 7 whorls), D: 44.1 mm; Fig. 6F, G • 1 spm; idem; CPAG.RAN.I.100 (cast MNHN. F.A93711), H: 98.2 mm (c. 23 whorls), D: 28.8 mm; Fig. 6D, E • 1 spm; idem; CPAG.RAN.I.101 (cast MNHN. F.A93724), H: 17.8 mm (shell incomplete, 3 whorls), D: 12.3 mm. OTHER EXAMINED MATERIAL. — Pakistan • 1 spm; same data as the holotype; MNHN. F.A93696 • 1 spm; idem; MNHN. F.A93697 • 1 spm; idem; MNHN. F.A93698 • 2 spms; idem; MNHN. F.A93699 • 1 spm; idem; MNHN. F.A93700 • 2 spms; idem; MNHN. F.A93701 • 3 spms; idem; MNHN. F.A93702 • 1 spm; idem; MNHN. F.A93704 • 5 spms; idem; MNHN. F.A93705 • 1 spm; idem; MNHN. F.A93706 • 3 spms; idem; MNHN. F.A93707 • 2 spms; idem; MNHN. F.A93708 • 1 spm; idem; MNHN. F.A93709 • 2 spms; idem; MNHN. F.A93710. ETYMOLOGY. — From the type formation: Lakhra Formation. TYPE HORIZON. — Diversified molluscs bed, base of the East-Lakhra village section (stn 2), lower Lakhra Formation, early Eocene (Ypresian). TYPE LOCALITY. — Pakistan, Sindh Province, Jamshoro district; Lakhra Dome, East-Lakhra village section (stn 2). DISTRIBUTION. — Lakhra Formation, early Eocene (Ypresian). DESCRIPTION Medium sized shell (H = 136 mm) of more than 33 teleoconch whorls starting with 24 turriculate whorls resembling a column, followed by eight whorls becoming increasingly convex; overall aspect of shell, turriculate, coleoconid. Antepenultimate and penultimate whorls slightly more inflated that earlier spire whorls. Apical angle 18-19°. Last whorl 29.3% total height (holotype). Whorls separated by narrowly impressed suture. Protoconch and first teleoconch whorls unknown. Delicate spiral sculpture present on early teleoconch whorls, disappearing on last whorls – Detailed composite description of the spiral sculpture: holotype MNHN.F.A93729 (Fig. 6A, B, H: 136.8 mm), primary cords P1-P3 present until 20th whorl, P1 close-set to the adapical suture more prominent than P2 and P3, P2 less prominent.On whorl no. 21, appearance of s2 between P2 and P3, P2 less prominent than the other primary cords. From whorl no. 22 to no. 24 (Fig. 6B), s2 becoming larger and subequal to P2, s2 and P2 less prominent that the other cords; small granules visible on cords. On whorls nos. 25-26 (Fig. 6B), P1, P2 and s2 starting to fade gradually, P3 still prominent. On whorl no. 27 (Fig. 6B), loss of spiral sculpture – Paratype MNHN.F.A93728 (Fig. 5A, H: 58.6 mm) on whorls no. 1 to no. 4, P1-P3 and s2 present; P1 prominent, close-set to adapical suture; P3 close-set to abapical suture; s2 close-set to P2 and less prominent. On whorls nos. 5-6, P1, P2 and s2 starting to fade gradually, P3 still prominent. On whorl no. 7, P1 tending to fuse with P2. On whorl 8, loss of the spiral sculpture – Paratype MNHN.F.A93727 (Fig. 5B, H: 29.2 mm), on whorls no. 1 to no. 3, P1-P3 and s2 present; P1 prominent, close-set to adapical suture; P3 close-set to abapical suture; s2 close-set to P2 and less prominent. On whorl nos. 4-5, P1, P2 and s2 starting to fade, P3 still prominent. On whorl no. 6, loss of the spiral sculpture – Paratype MNHN.F.A93725 (Fig. 5C, H: 57.1 mm), on whorls no. 1 to no. 3, P1-P3 and s2 present; P1 prominent, close-set to adapical suture; P3 prominent close-set to abapical suture; P2 and s2 weaker; small granules on P2, s2 and P3. On whorl no. 4, P1, P2 to tending to fuse and s2 starting to fade gradually, P3 still prominent. On whorl no. 5, only P3. On whorl no. 6, loss of the spiral sculpture – Paratype MNHN.F.A93726 (Fig. 5D, H: 31.9 mm), on whorls nos. 1-2, P1-P3 and s2 present; P1 close-set to adapical suture; P3 close-set to abapical suture; s2 close to P2 and less prominent. On whorl no. 3, P1 and P2 tending to fuse. On whorl no. 4, only P3. On whorl no. 5, total loss of the spiral sculpture – No axial sculpture, except some growing striae. Aperture narrow, ovoid, compressed adapically, of 88.7% of diameter and 84.33% of length of last whorl. Anal canal short and narrow. Columella weakly concave with one fold; columellar lip slightly erect; parietal lip adherent. Siphonal canal moderately short, bent, turning left and dorsally curved. Outer lip prosocyrt, not thickened. Remains of periostracum imprints (paratype MNHN.F. A93711). Residual colour pattern consisting of red stripes, approximately parallel, with inclination varying from slightly sinuous to sigmoidal (paratype MNHN.F. A98132, Fig. 6F, G).