Nemesia eira Rafael & Molero-Baltanás, 2026, n. sp.

Nemesia eira n. sp. (Figs 4A; 5A; 6A, B; 7A, B; 8A, C) urn:lsid:zoobank.org:act: 6A71B871-850B-4EB4-993D-33295DE34830 TYPE MATERIAL. — Holotype. Spain • ♂; La Cañada de Los Ratones, Alcalá de los Gazules, Cádiz; 36°24’5”N, 5°44’14”W; Alt.: 53 m; 9.X.2019; MNCN 20.02/41854. Paratypes. Spain •, ♀; La Cañada de Los Ratones, Alcalá de los Gazules; Cádiz; 36°24’5”N, 5°44’14”W; Alt.: 53 m; 23.XI.2019; UCO – 052b • ♀; La Cañada de Los Ratones, Alcalá de los Gazules, Cádiz; 36°24’5”N, 5°44’14”W; Alt.: 53 m; 30.IV.2018; MNCN 20.02/41855 • ♀; La Cañada de Los Ratones, Alcalá de los Gazules, Cádiz; 36°24’5”N, 5°44’14”W; Alt.: 53 m; 3.XI.2018; UCO – 033b • ♀; La Cañada de Los Ratones, Alcalá de los Gazules, Cádiz; 36°24’5”N, 5°44’14”W; Alt.: 53 m; 20.IV.2019; UCO – 075d. ADDITIONAL MATERIAL EXAMINED. — Spain • ♂; La Cañada de Los Ratones, Alcalá de los Gazules, Cádiz; 36°24’5”N, 5°44’14”W; Alt.: 53 m; 23.IX.2019; Coll. CPP- 0001 • ♀; La Cañada de Los Ratones, Alcalá de los Gazules, Cádiz; 36°24’5”N, 5°44’14”W; Alt.: 53 m; 19.X.2019; Coll. CPP- 0003 • ♀; La Cañada de Los Ratones, Alcalá de los Gazules, Cádiz; 36°24’5”N, 5°44’14”W; Alt.: 53 m; 23.I.2022; Coll. CPP- 0195 • ♀; Pinar del Rey, San Roque, Cádiz; 36°15’0”N, 5°23’39”W; Alt.: 54 m; XII.2018; Coll. CPP- 0010 • ♀; Pinar del Rey, San Roque, Cádiz; 36°15’0”N, 5°23’39”W; Alt.: 54 m; 15.XII.2019; Coll. CPP- 0006 • ♀; Pinar del Rey, San Roque, Cádiz; 36°15’0”N, 5°23’39”W; Alt.: 54 m; 15.XII.2019; Coll. CPP- 0007 • ♀; Conil de la Frontera, Cádiz; 36°16’10”N, 6°0’45”W; Alt.: 111 m; 11.XII.2021; Coll. CPP- 0193 • ♂; Aguadulce, Rota, Cádiz; 36°40’3”N, 6°24’11”W; Alt.: 7 m; 24.X.2023; Coll. CPP- 0531. ETYMOLOGY. — “ eira ” is a noun in apposition and corresponds to the acronym formed by the initials of the full name of Ester Irene Reina Alfonso. This species is named in her honour, in recognition of her wholehearted companionship to the first author. DIAGNOSIS. — The palpal bulbs of males resemble those of N. shenlongi, although the tip is proportionally thinner and more elongated (Fig. 4A). Females of N. eira n. sp. have larger spermathecal receptacles than those of N. shenlongi, with a narrower distance between the spermathecae (Fig. 5A). The in vivo colour of females and immature specimens of N. eira n. sp. is yellowish orange, characterised by a distinctive dark mark between the posterior edge of carapace and the fovea (Fig. 6A; 7A). Males are orange to slightly greyish, displaying the same mark on the prosoma (Fig. 7B). Additionally, the shape of sternal sigillae differs between these species: in N. shenlongi they are circular (Pertegal et al. 2022. Fig. 4C), whereas in N. eira n. sp., they are ovoid (Fig. 6B). The distinctive colouration and markings on the carapace are characteristic in N. eira n. sp., enabling its differentiation from both the previously described species and those presented below. Nemesia eira n. sp. shares its distribution in some areas with N. amicitia (Pertegal & Molero-Baltanás, 2022), which can be distinguished from the new species by its compact, spherical bulbs with a curved and narrowed embolus in males, and by the funnel-shaped spermathecae with slightly curved tubes in females. Nemesia amicitia is smaller and darker in colour than N. eira n. sp., and their trapdoor types also differ. Nemesia amicitia constructs a rigid, horizontal wafer-type trapdoor that rests against the inner edge of a curb made of particles and silk surrounding the entrance, whereas the trapdoor built by N. eira n. sp. consists of a thin and flexible lid. DESCRIPTION Holotype: MNCN 20.02 /41854 Male General appearance. Brownish-orange prosoma with dark mark on the posterior area. Opisthosoma orange with slightly dark cardiac mark (Fig. 7B). Prosoma. Dark clypeus with five bristles and white pubescence. POP black, connecting all eyes. Caput low. Crest zone orange with seven short bristles.Shallow fovea.Chelicerae dark brown with whitish pubescence. Cheliceral rastellum with six strong bristles (sometimes seven or eight). Fang ridge smooth. Cheliceral furrow teeth with six marginal denticles. Sternum yellowish with poorly visible ovoid sigillae; the first two pairs of sigillae marginal, the third pair submarginal. Maxillae yellowish with two to three spiny cuspules, but other males with four. Labium and coxae yellow. Palp and legs I-IV. Cymbium dorsal patch with 20-23 spines on the holotype. Bulb globular, embolus with elongated duckbill-shaped end (Fig. 3A). Palp tibia ratio 2.33±0.18. Apical dorsal patch of tibia of palp with nine spines. Clasper hook slightly curved. Clasper field with scopula. Metatarsus I slightly curved. Femora of palp and legs brownish, remaining segments yellow. Scopula from the distal zone of tibiae to tarsi on legs I and II and only on the tarsi on the posterior pair of legs. MTF4 ratio: tibia> metatarsus> femur. Spines, Palp. Cy= 23:20 (16-17:18-19), Pach= 9:9 (12-15:10- 13); prolateral side, Pa: 2:2 (1:1); Leg I: prolateral side, Pa= 2:2, Ti= 4:3 (3:3), Met= 3:2 (1-3:2-3); retrolateral side, Pa= 1:1, Ti= 3:3, Met= 3:2 (1:2); Leg II: prolateral side, Pa= 2:2, Tib= 4:3, Met= 3:3; retrolateral side, Pa= 1:1 (0:0), Tib= 3:3 (2:3), Met= 3:2 (3:3); Leg III: prolateral side, Pat= 2:2, Tib= 3:4 (4:3), Met= 3:3; retrolateral side, Pa= 1:1, Tib= 3:3, Met= 3:3; Leg IV, prolateral side, Pa= 3:3 (2:2); Tib= 6.8 (5-6:6-7); Met= 3.4 (4:3); retrolateral side, Pat= 1:1; Tib= 4.4; Met= 5.5 (4-6:4). Opisthosoma. Epiandrous fusillae: 49; sub-pentagonal epiandric area: 0.82 mm 2; density: 59.76 epiandrous fusillae/ mm2. Other males, epiandrous fusillae: 45±3; epiandric area: 0.65± 0.14 mm 2; density: 70.82±9.83 epiandrous fusillae/ mm 2. Spinnerets: PLS: proximal segment as long as middle and distal segments together. PMS cone-shaped. Measurements (n = 3). Bl= 16.85 (10.12-16.85); Prosoma. Ca= 4.47 (3.96-4.47); CL= 7.63 (6.8-7.63); CW= 5.91 (5.49- 5.91); Ch= 1.99 (1.59-2.32); Th= 1.52 (1.23-2.2); Cly2= 0.21 (0.11-0.21); AER= 1.1 (1.05-1.13); PER= 1.18 (1.13-1.19); ALE= 0.32 (0.29-0.32); PLE= 0.29 (0.24-0.29); AME= 0.24 (0.2-0.33); PME= 0.24 (0.23-0.24); l= 0.58 (0.58-0.63); mal= 2.07 (2.07-2.11); maw= 1.09 (1.09-1.21); lal= 0.61 (0.57- 0.61); law= 1.02 (0.81-1.02); stl= 3.6 (3.23-3.6); stw= 2.87 (2.57-2.87); Palp: E1= 0.6 (0.52-0.6); E2= 0.2 (0.17-0.2); E3= 0.51 (0.39-0.51); Bu= 1.68 (1.54-1.68); Em= 0.76 (0.76-0.83); Tiw= 1.37 (1.11-1.37); Cyl= 1.37 (1.28-1.42); Til= 2.93 (2.63- 2.93); Pal= 1.81 (1.65-1.82); Fel= 3.64 (3.05-3.64); TOTAL= 9.75 (8.61-9.75); Leg I: Tal= 3.15 (3.15-3.32); Mel= 4.57 (4.07-4.57); Til= 4.21 (4.13-4.21); Pal= 3.79 (3.19-3.79); Fel= 6.33 (5.47-6.33); TOTAL= 22.05 (20.18-22.05); Leg II: Tal= 2.81 (2.78-2.81); Mel= 4.18 (4.01-4.18); Til= 3.78 (3.78-3.97); Pal= 3.36 (2.93-3.36); Fel= 5.92 (5.31-5.92); TOTAL= 20.05 (19.02-20.05); Leg III:Tal= 3.28 (2.95-3.28); Mel= 4.81 (4.76-4.97);Til= 3.76 (3.41-3.76); Pal= 2.72 (2.39- 2.72); Fel= 5.16 (4.3-5.16); TOTAL= 19.73 (17.86-19.73); Leg IV: Tal= 3.31 (3.16-3.31); Mel= 7.02 (6.54-7.02); Til= 7.15 (6.66-7.15); Pal= 3.81 (3.21-3.81); Fel= 6.6 (5.78-6.6); TOTAL= 27.89 (25.46-27.89). Paratype: MNCN 20.02/41855 Female General appearance. Orange prosoma with dark mark between fovea and posterior area. Opisthosoma greyish-orange with slightly dark cardiac mark and four chevrons (Fig. 7A). Prosoma. Dark clypeus with five bristles. POP in three groups covering lateral and middle eyes (Fig. 6A). Caput slightly elevated. Crest zone orange with seven bristles. Shallow fovea. Chelicerae dark brown (Fig. 6A). Cheliceral rastellum with six strong bristles (sometimes seven or eight). Fang ridge smooth. Cheliceral furrow teeth with seven to nine marginal denticles. Sternum yellowish with brown ovoid sigillae arranged as in holotype (Fig. 6B). Maxillae yellowish with six finger-shaped cuspules, 7-12 in other specimens. Labium and coxae yellow (Fig. 6B). Palp and legs I-IV. Yellow with tarsi and metatarsi darkened by scopulae. MTF4 ratio: tibia> femur> metatarsus, usually tibia> metatarsus> femur. Spines, Palp. Prolateral side, Pat= 2:2, Tib= 4:3 (2:2); retrolateral side, Tib= 1:1 (2:1); Leg I: prolateral side, Pa= 2:3, Tib: 3:3 (2:2); Leg II: prolateral side, Pa= 2:2, Tib= 3:3 (1-2:2), Met= 1:1 (2:1); Leg III: prolateral side, Pa= 2:1 (2:1), Tib= 2:2 (2:3), Met= 3:3 (2:2); retrolateral side, Tib= 2:4 (2-3:2- 3), Met= 3:4 (3:3); Leg IV: prolateral side, Met= 2:2 (1:2); retrolateral side, Tib= 2:2 (2:3), Met= 3:3 (2:3). Opisthosoma. Spinnerets: PLS: proximal segment as long as middle and distal segments together. Maculae absent. PMS: digitiform. Spermatheca: elongated, bag-shaped, with receptacles close together at their bases (Fig. 4A). Measurements (n = 10). Bl= 19.58 (16.37-19.58); Prosoma. Ca= 5.59 (4.53-5.59); CL= 9.62 (7.19-9.62); CW= 7.11 (5.8- 7.36); Ch= 3.45 (2.4-3.45); Th= 2.19 (1.19-2.2); Cly2= 0.26 (0.26-0.4); AER= 1.4 (1.17-1.53); PER= 1.54 (1.26-1.62); ALE= 0.36 (0-0.4); PLE= 0.39 (0.31-0.44); AME= 0.23 (0.18-0.23); PME= 0.24 (0.11-1.56); l= 0.66 (0.63-0.75); mal= 3.15 (2.62-3.19); maw= 1.81 (1.47-1.85); lal= 0.86 (0.73-1.02); law= 1.58 (1.29-1.7); stl= 4.77 (3.81-4.77); stw= 3.45 (0.53-4); Palp: Tal= 2.91 (2.49-3.09); Til= 2.93 (2.77- 3.22); Pal= 2.53 (2.23-2.78); Fel= 4.66 (4.02-5.13); TOTAL= 13.03 (11.75-13.89); Leg I: Tal= 2.17 (1.77-2.47); Mel= 3.16 (2.97-3.74); Til= 3.84 (3.75-4.42); Pal= 4.12 (3.55-4.44); Fel= 6.67 (5.32-6.84); TOTAL= 19.96 (17.88-21.38); Leg II: Tal= 2.16 (1.69-2.16); Mel= 2.72 (2.52-3.67); Til= 3.64 (3.05-3.99); Pal= 3.52 (3.24-3.94); Fel= 5.73 (4.94-6.02); TOTAL= 17.77 (15.79-19.4); Leg III: Tal= 2.18 (1.77-2.59); Mel= 3.55 (2.81-3.71); Til= 2.95 (2.47-3.02); Pal= 3.1 (2.51- 3.23); Fel= 4.75 (3.77-4.96); TOTAL= 16.53 (13.77-16.95); Leg IV: Tal= 2.98 (2.06-2.98); Mel= 5.28 (4.36-5.7); Til= 7.1 (6.28-7.25); Pal= 4.6 (3.61-4.75); Fel= 6.18 (5.53-6.42); TOTAL= 26.14 (22.79-26.52). INTRASPECIFIC VARIATIONS It is occasionally possible to find some specimens exhibiting a slightly pinki