Staphylus (Capilla) buena

Staphylus (Capilla) buena (Williams & Bell, 1940) Figures 2 E, 2 F, 3 E, 3 F, 11 Pholisora buena Williams & Bell, 1940. Trans. Amer. Ent. Soc. 66: 131, fig. 12 (male gen.); holotype male, Buenavista, 75 km, northwest of Santa Cruz, Bolivia; ANSP. – Hayward, 1951. Rev. Soc. ent. arg. 15: 55. Staphylus buena; Evans, 1953. Cat. Amer. Hesp. 3, p. 86, pl. 37 (male gen.). – K. Brown & O. Mielke, 1967. Jour. Lep. Soc. 21 (3): 164. – Hayward, 1973. Op. Lill. 23: 43. – Biezanko & O. Mielke, 1973. Acta biol. paranaense 2: 69. – Bridges, 1983. Lep. Hesp. 1, p. 19; 2, p. 34. – K. Brown, 1987. An. Prim. Simp. Pantanal, Brasília, p. 167. – Bridges, 1988. Cat. Hesp. 1, p. 30; 2, p. 57. – Bridges, 1994. Cat. Fam. - Group, Gen. - Group, Sp. Group Nam. Hesp. (Lep.) World 8, p. 36; 9, p. 64. – O. Mielke, 2004. Hesperioidea, p. 55, in Lamas (ed.). Checklist: Part 4 A, Hesperioidea - Papilionoidea, in Heppner (ed.). Atlas Neotrop. Lep. 5 A. – O. Mielke, 2005. Cat. Amer. Hesperioidea 3, p. 722. – Gareca et al., 2006, in Gareca & Reichle. Marip. diurn. Bolivia, p. 18. – Palo Jr., 2017. Borb. Brasil 3, p. 2306. – Klimaitis et al., 2018. Marip. Argentinas, p. 233, figs. (male d, v, map). (no genus) buena; Beattie, 1976. Rhop. Direct., p. 100. Taxonomic history. Williams and Bell (1940) described Pholisora buena. Evans (1953) cited it as Staphylus buena, as have most subsequent authors that mentioned this species in taxonomic comments, faunistic studies and catalogues. Type. Pholisora buena Williams & Bell, 1940 was described from the male holotype from Buenavista, 75 km northwest of Santa Cruz, Bolivia, deposited in ANSP, currently in CMNH, with the following labels: / Type 7188 BUENA Will-Bell / Buenavista 75 Km. N. W. of Santa Cruz, Bolivia / Pholisora buena Williams + Bell Holotype ♂ / 450 m alt. 1926–27 (Jose Steinbach) / Genitalia slide No. 2294 /. Diagnosis. Staphylus (C.) buena is readily distinguished from other species of Staphylus (Capilla) by a combination of characters of the male genitalia (Fig. 3 E, F): sacculus bearing a dense set of small spines internally and ampulla produced at distodorsal margin, bearing a dense set of small, socketed spines, proximally concave with a very produced dentate area on the ventral-internal margin. Male description. Fig. 2 E, F. Head: Brown with golden scales dorsally; presumably yellow ventrally. FW length: 1.3 cm. HW length: 1 cm. DFW: Brown, two subapical white dots. VFW: Same white markings as on DFW. VHW. Brown. Genitalia (Fig. 3 E, F). Dorsal view of uncus unknown, apparently conical, curving downwards in lateral view, tapering distally, bearing a tuft of short hair-like scales at the base. Valva: sacculus scalene, triangular-shaped, bearing a dense set of small spines internally; harpe rounded at distal margin, not exceeding ampulla; ampulla broad, produced at distodorsal margin, bearing a dense set of small, socketed spines, proximally concave with a very produced dentate area on the ventral-internal margin. Aedeagus approximately the same length of valva, bearing small denticles on the distal portion. Fultura superior bearing two dense clusters of apparently large, stout spines laterally. Female description. Unknown. Distribution. Fig. 11. Bolivia: Buena Vista; Argentina: Salta (Hayward 1973), Santa Fé (Hayward 1973) and Tucumán (Hayward 1973). Temporal distribution. Unknown. Etymology. Although not mentioned in the description, the epithet of Staphylus (C.) buena clearly is in reference to the type locality, Buenavista (Bolivia). Remarks. As the holotype genitalia was not studied, all the morphological interpretations are from the drawing of the genitalia in the original description (Fig. 3 E). In the original drawing, the dorsal view of the genitalia is not illustrated, also there is no indication of the saccus. It is assumed that the saccus is triangular and broad, as the other species of the subgenus. In the costa region of the valva, there is a set of spines. However, the species of Staphylus (Capilla) do not have a well-defined costa. These species usually have a fultura superior developed as a plate bearing two dense clusters of small stout spines laterally. Therefore, in this study, the spines represented in the costa region of the original drawing are treated as the spines of the fultura superior.