Trichaphodiellus brasiliensis

Trichaphodiellus brasiliensis (Castelnau, 1840) Figs 4D–F, 38A Aphodius brasiliensis Castelnau, 1840: 65. Aphodius excellens Endrödi, 1964: 187. Aphodius (Trichaphodiellus) brasiliensis – Schmidt 1913: 167. Trichaphodiellus brasiliensis – Dellacasa et al. 2001a: 288. Diagnosis Same as for the genus. Material examined BRAZIL – Acre • 2 specs; Rio Branco; CEMT • 1 spec.; same data as for preceding; 10°01′58.09″ S, 67°42′12.74″ W; 9 Aug. 2015; A. Specht leg.; light; CEMT. – Amazonas • 1 spec.; Presidente Figueiredo, Ilha Pantanal; 1°47′49″ S, 59°26′53″ W; 22–24 Jun. 2016; L. Sawaris leg.; submontane dense ombrophilous forest; human and pig dung; CEMT. – Espírito Santo • 1 spec.; Linhares, AFCVRD; Oct. 1988; J.S. Santos leg.; CEMT • 2 specs; same data as for preceding; Reserva Natural Vale; 19°09′04″ S, 40°04′14″ W; Dec. 1987; J.S. Santos leg.; light; CEMT. – Mato Grosso • 1 spec.; Alta Floresta, CEPLAC; 4 Sep. 2009; V. Gonçalves leg.; Tapirus feces; CEMT • 1 spec.; same data as for preceding; CEMT • 3 specs; same data as for preceding; CEMT • 1 spec.; Apiacás, Parna Juruena; 10 Mar. 2011; M. Lima leg.; CEMT • 1 spec.; Chapada dos Guimarães, Santuário dos Elefantes; 15°18′50″ S, 55°24′19″ W; 21 Jan. 2020; H. Sehn et al. leg.; elephant feces; CEMT • 4 specs; same data as for preceding; 15°18′55″ S, 55°24′14″ W; 22 Jan. 2020; CEMT • 1 spec.; same data as for preceding; 15°18′22″ S, 55°24′14″ W; 23 Jan. 2020; CEMT • 1 spec.; same data as for preceding; 15°18′55″ S, 55°24′07″ W; CEMT • 1 spec.; same data as for preceding; 15°18′50″ S, 55°24′12″ W; 24 Jan. 2020; CEMT • 3 specs; Poconé; 26 Apr. 1998; light; CEMT • 1 spec.; Corumbá, Porto Jofre, Fazenda São Bento; 13 Apr. 2011; R.V. Nunes leg.; CEMT • 1 spec.; Cotriguaçú, Fazenda São Nicolau; 10 Dec. 2009; M.R. Barreto leg.; forest; CEMT • 1 spec.; Poconé, Base UFMT; 16°29′56″ S, 56°24′49″ W; Mar. 2012; M. Rossini leg.; CEMT. – Mato Grosso do Sul • 1 spec.; Chapadão do Sul; 18°46′30.09″ S, 52°31′04.98″ W; 5 Jan. 2016; A. Specht leg.; light; CEMT. – Minas Gerais • 1 spec.; Conceição do Mato Dentro, 23K; 67°69′53″ S, 79°11′57″ W; elev. 711 m; 16 Sep. 2012; Uceli and Borges leg.; CEMT • 1 spec.; same data as for preceding; 66°31′39″ S, 79°11′235″ W; elev. 704 m; 20 Sep. 2012; Uceli and Borges leg.; light; CEMT • 1 spec.; Cordisburgo; Jul. 1992; F. Zagury leg.; CEMT • 1 spec.; same data as for preceding; Fazenda Pontinha; Jan. 1996; Vaz-de-Mello leg.; CEMT • 1 spec.; Marlieria; Jul. 1994; Mello leg.; CEMT • 1 spec.; Viçosa; 11 Feb. 1996; J.N.C. Louzada leg.; CEMT • 4 specs; Pains, ISLA G. do Zé Serafim III; 25 Aug. 2009; R.A. Zampauto leg.; CEMT • 6 specs; Paraopeba; Mar. 1997; S. Lourenço jr leg.; CEMT. – Pará • 1 spec.; Parauapebas; 5°56′46″ S, 50°12′17″ W; 26 Feb. 2019; F. França leg.; secondary forest; human and pig dung; CEMT. – Paraná • 2 specs; Guaratuba, Estrada dos Castelhanos; elev. 650 m; 25 Nov. 2009; Grossi et al. leg.; light; CEMT. – Paraíba • 1 spec.; Areia, Mata de Pau Ferro; 2 May–1 Jun. 2000; Avany and Maria leg.; light; CEMT. – Rio de Janeiro • 9 specs; Cachoeira de Macacu, Guapiaçu ER; 22°27′5″ S, 42°46′23″ W; elev. 33 m; 22 Sep. 2019; L.R. Martínez leg.; Tapirus feces; CEMT • 1 spec.; same data as for preceding; 25 Oct. 2019; horse feces • 7 specs; same data as for preceding; 10 Jan. 2020; CEMT • 1 spec.; Miguel Pereira; Mar. 1998; J. Carlos leg.; CEMT • 2 specs; Nova Friburgo; Nov. 1993; E.J. Grossi leg.; CEMT • 1 spec.; same data as for preceding; Dec. 1995; P. Grossi and E. Grossi leg.; CEMT • 1 spec.; same data as for preceding; Nov. 1997; CEMT • 2 specs; same data as for preceding; Macaé de Cima; Nov. 1998; F.Z. Vaz-de-Mello and E. Grossi leg.; CEMT • 1 spec.; same data as for preceding; Dec. 2000; P. Grossi leg.; CEMT • 1 spec.; Santa Maria Madalena, Pousada Verbicaro, RPPN, MAD 03; 29 Oct.–4 Nov. 2015; Mermudes and Mattos leg.; Pennsylvania trap; CEMT. – São Paulo • 1 spec.; São Miguel Arcanjo, P.E. Carlos Botelho; 24°03′58″ S, 47°59′53″ W; elev. 743 m; 6 Mar. 2012; M. Boutefeu leg.; Tapirus feces; CEMT • 1 spec.; same data as for preceding; 24°03′59″ S, 47°59′39″ W; elev. 580 m; 8 Apr. 2012; CEMT • 1 spec.; same data as for preceding; 4°40′01″ S, 47°59′40″ W; elev. 595 m; 9 Apr. 2012; CEMT • 1 spec.; same data as for preceding; 24°03′50″ S, 47°59′23″ W; elev. 747 m; CEMT • 4 specs; same data as for preceding; 11 Jun. 2012; CEMT • 1 spec.; same data as for preceding; 13 Jun. 2012; CEMT • 1 spec.; same data as for preceding; 15 May 2012; CEMT. – Sergipe • 2 specs; São Cristóvão; 24 Aug. 2016; M.F. Santos leg.; CEMT. COLOMBIA – Antioquia • 3 specs; Río Negro, Capiro; Abr. 1995; F. Vallejo leg.; CEMT. – Cundinamarca • 1 spec.; La Calera; Jul. 2000; J. Noriega leg.; CEMT • 1 spec.; Bogotá; Sep. 1996; J. Noriega leg.; CEMT • 1 spec.; same data as for preceding; Sep. 1998; CEMT • 1 spec.; Nocaima; 5°04′16″ N, 74°22′61″ W; elev. 1100 m; Sep. 2005; J. Noriega leg.; culture of sugar cane; CEMT. – Meta • 1 spec.; San. Martín, Tocancipa; elev. 330 m; Abr. 2006; P. López leg.; sabama forest; CEMT • 1 spec.; same data as for preceding; N. Montoya leg.; pitfall; CEMT. ECUADOR – Chimborazo • 1 spec.; Huigra, Chimborazo; elev. 1850 m; 31 Dec. 1997; G. Onore leg.; CEMT • 3 specs; same data as for preceding; 1 Jan. 1998; CEMT. – El Oro • 23 specs; Piñas; elev. 1250 m; 1 Sep. 1997; C. Carpio leg.; cow feces; CEMT • 5 specs; same data as for preceding; 2 Sep. 1997; human feces CEMT. – Loja • 1 spec.; Río Catamayo; elev. 1500 m; 29 Aug. 1997; C. Carpio leg.; CEMT. – Morona-Santiago • 2 specs; Bosque Domono; elev. 1650 m; 22 Jul. 1997; C. Carpio leg.; CEMT • 1 spec.; Vía Plan Milagro-Gral Gutiérrez Km 5; elev. 1500 m; 19 Aug. 1997; C. Carpio leg.; CEMT • 4 specs; same data as for preceding; feces; CEMT • 3 specs; Yunganza; elev. 1500 m; 28 Aug. 1997; C. Carpio leg.; cow feces; CEMT • 1 spec.; Palmeras; elev. 1800 m; 19 Nov. 1994; P. Guardenas leg.; forest; CEMT. – Napo • 4 specs; Cosanga; 0°35′36″ S, 77°56′18″ W; elev. 1750 m; 25 Mar. 1996; T. Jaramillo leg.; CEMT • 1 spec.; Vía Hollín-Loreto Km 3; elev. 1100 m; 6 Dec. 1987; A. Cordova leg.; CEMT • 1 spec.; same data as for preceding; Km 25; elev. 1100 m; P. Mendoza leg.; CEMT • 2 specs; same data as for preceding, Km 3; A. Cordova leg.; CEMT • 1 spec.; same data as for preceding; Km 60; R. Figuerda leg.; CEMT • 1 spec.; same data as for preceding; Km 17; M. Peñaherrera leg.; CEMT. – Pichincha • 3 specs; Alangasi; 19 Jan. 1988; M.E. Sánchez leg.; CEMT • 2 specs; El Tingo; 5 May 1997; P. Leone leg.; CEMT • 1 spec.; same data as for preceding; 30 Jan. 1988; CEMT • 1 spec.; Mindo; elev. 1500 m; 30 Dec. 1994; P. Salazar leg.; CEMT • 2 specs; same data as for preceding; elev. 1400 m; 8 Jun. 1995; F. Sancho leg.; CEMT • 1 spec.; Quito, Alangasi; 19 Jan. 1988; P. Merdoxa leg.; CEMT • 1 spec.; Río Silanchi; elev. 640 m; 30 Apr. 1991; E. Baus leg.; CEMT • 5 specs; same data as for preceding; Mindo; elev. 1200 m; 17 Jun. 1995; M. Lascano leg.; CEMT • 3 specs; same data as for preceding; elev. 1250 m; 8 Jul. 1995; M. Vallejo leg.; CEMT • 1 spec.; same data as for preceding; 10 Jul. 1995; CEMT • 1 spec.; Pam. Argentin; elev. 1500 m; Apr. 1988; M. Griialua leg.; CEMT. – Zamora • 6 specs; Ch Las Peñas; elev. 1000 m; 24 Aug. 1997; C. Carpio leg.; CEMT. PERU – Madre de Dios • 1 spec.; 6 km NE of Mazuko; elev. 380 m; 19 Aug. 2012; Cavichioli et al. leg.; CEMT. VENEZUELA – Mérida • 2 specs; Bailadores; Jan. 1984; B. Joffre leg.; CEMT. Sexual dimorphism Males with clypeal tubercle larger than that of female; females with metaventrite plate nearly flat. Collecting information This species can be collected with light traps and is attracted to equine, bovine, Tapirus and human feces (Mesquita et al. 2017). Specimens were collected at altitudes ranging from 33 to 1850 metres.