Ambigolimax waterstoni Hutchinson, Reise & Schlitt 2022

Ambigolimax waterstoni Hutchinson, Reise & Schlitt, 2022 Figs 17–20, 69 Limax nyctelius – Connolly 1939: 176 –177 – Algeria, Morocco – not Bourguignat 1861. Quick 1960: 200 –201, fig. 17 B, C – Shebin El Kom, Egypt; Algeria – not Bourguignat 1861. Limax (Limacus) nyctelius – Regteren Altena 1966: 190 –192 – Algeria – not Bourguignat 1861 Lehmannia nyctelia – Wiktor 1983 a: 161 –162 – Algeria: Hamma. Jardin d’Essai; Lakhdaria (= Gorges de Palestro), rocky gape of the river Isser; Defile Sebaou near Tizi Ouzou Kabylia; Kabylia, mountains halfway between Mirabeau and Dra-el-Mizan – not Bourguignat 1861 Ambigolimax waterstoni – Hutchinson et al. 2022: 38 –40, figs 5, 6, 12 B, C. Material examined. • 2 exx; Boumerdès, Corso; 36°45.983'N, 3°26.817'E; 10 m a. s. l.; January 2019; R. Ramdini leg. • 2 exx; Tizi-Ouzou, Akvil, Ait Ouavan forest; 36°29.017'N, 4°17.000'E; 855 m a. s. l.; February 2022; R. Ramdini leg. Diagnosis. Body slender, length ≤ 55 mm; mantle and keel like the previous species, brown to yellowish with two dark bands on the mantle and posterior body sides; occasionally, there is an additional, more or less clear band along the middle of the mantle (Figs 17, 18). Genitalia characterised by a long (4–6 mm) cylindrical penis, often swollen at the apex; the short vas deferens and the penial retractor muscle enters near penial apex beside one another; duct of the bursa copulatrix arising from the distal penis (Fig. 19); inside the penis two flaps originate from the penial apex and run parallel, decreasing in height towards the atrium (Fig. 20) (Connolly 1939; Quick 1960; Giusti 1976; Barker 1979; Wiktor 1983 a; Hutchinson et al. 2022). Distribution. Ambigolimax waterstoni is presumed native to North Africa, but it is known as an alien species from Scottish greenhouses, Elba Island (Italy), the USA, South Africa, Australia, and New Zealand (Herbert 2010; Hutchinson et al. 2022). Ecology. Found in humid places, under stones, it can be synanthropic. Remarks. This species has been accepted for many decades under the name Lehmannia nyctelia or Ambigolimax nyctelius; however, Hutchinson et al. (2022) proposed the taxon nyctelia Bourguignat, 1861 as not available for Lehmannia or Ambigolimax. Indeed, the description and illustration of the pneumostome in the anterior part of the mantle (Bourguignat 1861) unambiguously refer nyctelius to the arionid genus Letourneuxia. Hutchinson et al. (2022) introduced the new name A. waterstoni for the North African native Ambigolimax species (previously named A. nyctelia) characterised by a long cylindrical penis, two parallel folds inside the penis, vas deferens, and penial retractor muscle entering near the penial apex.