Rheocricotopus (Psilocricotopus) chinogodavarius Niitsuma & Tang, 2026, sp. nov.
Abstract
Rheocricotopus (P.) chinogodavarius sp. nov. urn:lsid:zoobank.org:act: A8CA2B84-409B-4D28-978A-AC16975F929A Fig. 17, Table 15 Diagnosis Male Abdomen bicoloured, with tergite I entirely yellow, tergites II–IV entirely or partly dark brown, tergites V–IX entirely dark brown. Legs largely dark brown with apex of tibia and base of basitarsus broadly whitish. Humeral pit large, oval. Wing cuneiform with a distinct costal extension. Fore LR very high (ca 0.9). Superior volsella plate-like, low. Inferior volsella triangular. Crista dorsalis triangular, more or less conspicuous, positioned at about middle of gonocoxite. Etymology From the type locality China, and Rheocricotopus (P.) godavarius Lehmann, the male of which somewhat resembles that of the present species. Type material Holotype CHINA – Yunnan • ♂; Jinping, Ma’andi, unnamed stream; 22°46′ N, 103°31′ E; 1360 m a.s.l.; 8 Jun. 2017; J. Chen leg.; JNU, JNU 170724006. Paratypes CHINA – Yunnan • 2 ♂♂; Lijiang, Yulong, Jiuhe, 99-lakes scenic spot, Lake Tiancai; 26°38′ N, 99°43′ E; 3900 m a.s.l.; 18 Aug. 2023; H.Q. Tang leg.; JNU. Description Male (n = 3) Total length 2.1–2.9, 2.5 mm. COLOURATION. Head dark brown on vertex. Thorax mostly dark brown except yellow pleural membrane and pale HP; scutal vittae indistinct. Abdomen (Fig. 17A–C) bicoloured. T I entirely yellow. T II mostly yellow with mesal longitudinal, and anterior and posterior transverse dark bands; or with posterior transverse dark band only. T III entirely dark brown; or yellow with mesal and lateral longitudinal, and anterior and posterior transverse dark bands. T IV yellow with mesal longitudinal, and anterior and posterior transverse dark bands; or mesal longitudinal short band only. T V–IX entirely dark brown. Legs (Fig. 17D–E) dark brown, with whitish bands on ti and ta 1; fe and Ta 2–5 entirely dark brown in all legs. P 1 whitish on apical ⅔–¾ of ti and basal ½ of ta 1. P 2 and P 3 somewhat paler on apical half of ti and basal half of ta 1. HEAD. Temporals 7–14, 9. AR 0.38 (1). Clypeus with 4–8, 6 setae. Lengths (μm) of Pm 1–5: 25–37, 32; 49–74, 62; 103–160, 134; 125–155, 140; 221–290, 257, respectively. Pm 4 /Pm 3 1.0–1.2, 1.1; Pm 5 /Pm 4 1.8–1.9, 1.8. Pm 3 with 4–9, 6 SCl on 2 oval depressions. THORAX. Lateral Aps 5–8, 6. Ac 7–17, 13; longest 15–17, 16 μm long; Dc 5–8, 6, including 1 H; Pa 3–4, 3; Scts 4 (2). HP (Fig. 17F) large, oval. WING (Fig. 17G). Length 1.6–2.7, 2.2 mm. C extended moderately beyond end of R 4+5, extension 57–98, 81 μm long. VR 1.0–1.1, 1.1. R with 4–10, 7 setae, R 1 and R 4+5 bare. Sq with 1–4, 2 setae. AnL much reduced. LEGS. P 1 with ti spur 34–37, 36 μm long; P 2 with 2 ti spurs 15–20, 18 and 15–17, 16 μm long; P 3 with 2 ti spurs 39–47, 43 and 17–20, 18 μm long. P 2 without Sch. P 3 with ti comb of 11–16, 13 bristles. Lengths and proportions of legs as in Table 15. ABDOMEN. Hypopygium (Fig. 17H) with AnP pointed apically, bearing 7–11, 9 lateral setae. Gc 153–204, 179 µm long, 3.2–3.9, 3.4 × as long as broad at middle. SVo thin, roundly protruded, IVo (Fig. 17I) triangular. Gs (Fig. 17J) 74–91, 84 µm long, nearly parallel-sided, apically truncated, with CD triangular, somewhat variable in size, more or less conspicuous, positioned at about middle. HR 2.1–2.2, 2.1. Female, pupa and larva Unknown. Distribution The species is known only from Yunnan Province in Oriental China. Remarks The male somewhat resembles that of R. (P.) godavarius Lehmann, 1969 from Nepal in the bicoloured abdomen, the low antennal ratio (AR less than 0.4), the wing with a distinct costal extension and a much-reduced anal lobe, the plate-like superior volsella, and distinct, triangular crista dorsalis, but differs from it in the large oval humeral pit, the wing with squamal setae, the bicoloured legs with broad whitish bands on the tibia and the basitarsus, and the distinct, occasionally very large, crista dorsalis located around the middle of gonostylus. According to Lehmann (1969: 361–362; 363 fig. 19c, 370 fig. 9, 372 fig. 14b), the latter has an indistinct small humeral pit, wings with a bare squama, evenly brownish to brown legs, and a smaller crista dorsalis on the subapex of gonostylus.
Fields of Study & Subjects
Diptera species taxonomy and behavior
Stonefly species taxonomy and ecology
Freshwater macroinvertebrate diversity and ecology
Publication details
- Journal / source
- Zenodo (CERN European Organization for Nuclear Research)
- Publication date
- 30 January 2026
- Language
- EN