Rheocricotopus (Psilocricotopus) protuberans Niitsuma & Tang, 2026, sp. nov.

Rheocricotopus (P.) protuberans sp. nov. urn:lsid:zoobank.org:act: BD992B4C-9BEF-4833-B876-FCFD5EE39E7C Fig. 23, Table 21 Diagnosis Male Abdomen and legs dark brown. AR near 1.0. Antepronotal setae divided in medial and lateral groups. Humeral pit large, oval. Wing with obtuse anal lobe, short costal extension and well-developed squamal fringe. Superior volsella roundly protruded, thin. Inferior volsella triangular, simple. Crista dorsalis very large, conspicuous, occupying apical ⅔ of gonostylus, obtuse or right-angled at apex. Etymology From the Latin ‘ protuberans ’ (‘protuberant’), referring to the conspicuous crista dorsalis. Type material Holotype CHINA – Xizang • ♂; Lhoka, Nagarze, Lake Yamdrok; 28°56′ N, 90°42′ E; 4427 m a.s.l.; 18 Jul. 2014; H.Q. Tang leg.; JNU, JNU 160206012. Paratype CHINA – Xizang • 1 ♂; same data as for holotype; JNU. Description Male (n = 2) Total length 2.9–3.1 mm. COLOURATION. Head dark brown on vertex. Body entirely dark brown except pale HP. Legs dark brown. HEAD. Temporals 2. AR 0.85–0.88. Clypeus with 11 setae. Lengths (μm) of Pm 1–5: 37–42, 59–69, 116– 128, 148–160, 204–246, respectively. Pm 4 /Pm 3 1.3, Pm 5 /Pm 4 1.4–1.5. Pm 3 with more than 10 SCl. THORAX. Lateral Aps 2, and medial Aps 3. Ac 10–12, longest 12–17 μm long; Dc 7–8, including 0–1 H; Pa 3; Scts at least 4. HP (Fig. 23A) large, oval. WING (Fig. 23B). Length 2.2–2.4 mm. C extended slightly beyond end of R 4+5, extension 25–44 μm long. VR 1.1. R with 5–6 setae, R 1 and R 4+5 bare. AnL obtuse. Sq with 15–18 setae. LEGS. P 1 with single ti spur 39–44 μm long; P 2 with 2 ti spurs 22 (1) and 15–20 μm long; P 3 with 2 ti spurs 47–49 and 17–20 μm long. P 2 without Sch. P 3 with ti comb of 13–14 bristles. Lengths and proportions of legs as in Table 21. ABDOMEN. Hypopygium (Fig. 23C) with AnP pointed apically, bearing 7–8 lateral setae. Gc 185–192 µm long, 2.7–2.8 × as long as broad at middle, with SVo roundly protruded; IVo triangular, simple. Gs 86–91 µm long; CD triangular, obtuse or right-angled at apex, very large, occupying apical ⅔ of Gs. HR 2.11–2.14. Female, pupa and larva Unknown. Distribution The species is known from the type locality, Xizang Autonomous Region in Oriental China. Remarks The male resembles that of R. (P.) serratus in the antennal ratio of nearly 1.0, the large oval humeral pit, and the wing with an obtuse anal lobe, a well-developed squamal fringe and a costal vein slightly extended, but differs from it in the antepronotal setae divided into medial and lateral groups, and the crista dorsalis conspicuously large, obtuse or right-angled at the apex, the shape of which is unique among the known males of this genus including the orientalis group with a large median crista dorsalis. In the latter, antepronotal setae are located laterally only, and the crista dorsalis is much smaller, sawtooth-shaped, acute-angled at the apex. The male is similar also to that of R. (P.) insularis Makarchenko & Makarchenko, 2005 in the antennal ratio close to 1.0, the antepronotal setae located medially and laterally, and the wing with an obtuse anal lobe, but separable from it by the wing with a slight costal extension (25–44 μm long) and the conspicuously large crista dorsalis. According to Makarchenko & Makarchenko (2005: 128 figs 6–7; 129), the latter has wings with a well-developed costal extension (96 μm long), and much-smaller crista dorsalis.