Additional Specimens and New Localities of the Matang Caecilian, Ichthyophis biangularis (Amphibia, Gymnophiona, Ichthyophiidae), with Notes on Genetic and Geographic Diversity and Natural History

Species of Ichthyophis, from East, Southeast, and South Asia, are difficult to identify, mainly because many species were described based only on a few specimens without examining intraspecific variation. We redescribe I. biangularis, an endemic species of Malaysian Borneo. The morphology of the newly collected topotype is consistent with the original description. The phallodeum and the larvae are first described in this study. We sequenced Cyt b of mitochondrial DNA of this topotype and genetically identify the specimens collected in adjacent areas to the type locality. Intraspecific genetic variation of the species is relatively high compared to another species in this genus. We also describe the natural history of this species, including its habitat and reproduction.