A new species of Asiaharpa (Neogastropoda: Harpidae) from the Miocene of West Java, Indonesia, with a reappraisal of harpid phylogeny

A new species of Harpidae, Asiaharpa javanensis n. sp., is described from a fossil site ca. 12 My old, near Cisadane river in Cisauk district, Tangerang Regency, West Java, Indonesia. The phylogenetic position of Asiaharpa is reevaluated through a reassessment of conchological characters; it is a further example of the extraordinary diversification of the shallow marine fauna in the Sunda shelf during the Middle Miocene. The subfamily Harpinae is divided into three tribes: the fossil Eocitharini n. trib. (including only Eocithara Fischer, 1883 from the Paleocene to the Oligocene), Austroharpini n. trib. (also monotypic: Austroharpa Finlay, 1931 since the Oligocene), and the nominotypical Harpini including the basal Plesioharpa n. gen. from the Oligocene, the Oligocene to Recent Harpa Röding, 1798, and Asiaharpa Raven, 2021 from the Miocene of Sunda.